City Customization items can change the appearance of your base. You can activate them by tapping your base on the World Map and choosing Customize.

City Skins
Once unlocked, these Customization items are permanently added to your collection. You can Activate and Deactivate a City skin at any time, as long as you have unlocked it. City skins provide not only a different visual for your whole base, but also give you some useful bonuses.
Bastion of Mercy - while Active, provides a 5% bonus to your Recovery Speed.
Bastion of Loyalty - while Active, provides a 5% bonus to your Reinforcement Capacity.
Bastion of Wrath - while Active, provides a 5% bonus to your Recruitment Speed.
HQ Customization
Their effect is temporary and expires after a certain time. You can check how much time remains until your active customization item expires.
Empire Skytower - The view from the top floor is second to none
Eiffel Tower - Was build in 1889, so handle with care
Saint Basil's Cathedral - The top dome shines brighter than the brightest star
Desert Palace - Elegance in perfect symmetry
Note that activating a Customization item while another one is active will override the previous item.